just a first reaction ...fusable link
This sounds like the culprit.
If you can crank it from the starter relay the battery connection is good to that point. The fusible link is located between the Starter relay and the bulkhead connector to the dash.
It looks like a wire but the OEM ones had a tag molded onto them that said Fusible link. The cover feels rubbery instead of like plastic.
When the fusible link burns the center of the wire fries and the outer cover stays intact. DO NOT REPLACE IT WITH A REGULAR WIRE!
Next step in the wiring that goes bad is the bulkhead connector where the fusible link plugs into.
Check it too.
Lastly look at the large red and black wires behind the fuse block. If they are burnt it's time for a rewire but I think B1MAXX is correct and it's the fusible link or the bulkhead connector.